In a muscle contraction, the force that shortens the sarcomeres comes from the myosin cross-bridges pulling on the actin filaments.
A myosin head essentially functions like a cocked spring that bends and delivers a power stroke when it binds to an actin filament. A single sarcomere is forced to contract and shortens as a result of the power stroke's actin filament sliding past the myosin.
If there is enough ATP and the myosin-binding sites are exposed, myosin binds to actin to start the cross-bridge cycling process. The muscle contracts as the sarcomere shortens. Since this binding cannot take place without calcium, free calcium is a crucial regulator of muscle contraction.
Therefore,In a muscle contraction, the force that shortens the sarcomeres comes from the myosin cross-bridges pulling on the actin filaments.
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